Matayba arborescens

<<<Calyx>Petals>Appendages>Hairs

sericeous-tomentose1

1. 006-006-005-001

<<<Calyx>Petals>Appendages>Shape

and is fused to the petal’s base to form a pocket2

2. 006-006-005-002

<<<Calyx>Stamens>Filaments>Hairs

tomentose for most of their length3

3. 006-008-003-001

<<<Leaves>Leaflets>Apex>Shape

obtuse or less often abruptly acuminate into an obtuse, retuse tip4

4. 003-003-007-001

<<<Leaves>Leaflets>Base>Shape

acute to obtuse, sometimes slightly asymmetrical5

5. 003-003-006-001

<<<Leaves>Leaflets>Margins>Margin type

crenate6

6. 003-003-008-001

<<Calyx>Disk>Hairs

puberulent7

7. 006-007-001

<<Calyx>Lobes>Length

0.4-0.5 mm long8

8. 006-005-002

<<Calyx>Lobes>Shape

triangular9

9. 006-005-001

<<Calyx>Ovary>Hairs

appressed-pubescent10

10. 006-009-001

<<Calyx>Petals>Appendages

with a single appendage that is twice as long as the petal,11

11. 006-006-005

<<Calyx>Petals>Colour

white12

12. 006-006-001

<<Calyx>Petals>Hairs

sparsely tomentose13

13. 006-006-004

<<Calyx>Petals>Length

ca. 0.5 mm long14

14. 006-006-003

<<Calyx>Petals>Shape

rhombate, clawed15

15. 006-006-002

<<Calyx>Stamens>Filaments

the filaments ;16

16. 006-008-003

<<Calyx>Stamens>Length

ca. 3 mm long17

17. 006-008-002

<<Calyx>Stamens>Stamen number

818

18. 006-008-001

<<Capsules>Endocarp>Hairs

glabrous19

19. 007-008-001

<<Capsules>Pericarp>Texture

coriaceous20

20. 007-007-001

<<Flowers>Pedicels>Length

ca. 1 mm long21

21. 005-001-001

<<Leaves>Leaflets>Apex

the apex ,22

22. 003-003-007

<<Leaves>Leaflets>Base

the base ,23

23. 003-003-006

<<Leaves>Leaflets>Dimensions

8-17 (23) × 4-6.2 (7) cm24

24. 003-003-005

<<Leaves>Leaflets>Hairs

glabrous25

25. 003-003-003

<<Leaves>Leaflets>Leaflet number

(4) 6 (8)26

26. 003-003-001

<<Leaves>Leaflets>Margins

the margins ;27

27. 003-003-008

<<Leaves>Leaflets>Shape

elliptic or oblong-elliptic28

28. 003-003-004

<<Leaves>Leaflets>Texture

chartaceous29

29. 003-003-002

<<Leaves>Petiole plus rachis>Length

(5.5) 7-10 (20) cm long30

30. 003-001-001

<<Leaves>Petiolule>Hairs

puberulent31

31. 003-002-002

<<Leaves>Petiolule>Length

4-7 mm long32

32. 003-002-001

<<Leaves>Petiolule>Shape

enlarged at base33

33. 003-002-003

<<Leaves>Veins>Colour

usually drying darker than the blade34

34. 003-004-001

<<Seeds>Arillodes>Colour

white35

35. 008-004-001

<<Thyrses>Axes>Hairs

tomentulose36

36. 004-004-001

<Calyx>Colour

green37

37. 006-001

<Calyx>Disk

disc ;38

38. 006-007

<Calyx>Hairs

tomentulose39

39. 006-002

<Calyx>Length

0.6-1 mm long40

40. 006-004

<Calyx>Lobes

the lobes , ;41

41. 006-005

<Calyx>Ovary

ovary .42

42. 006-009

<Calyx>Petals

petals , , , , ;43

43. 006-006

<Calyx>Shape

cup-shaped44

44. 006-003

<Calyx>Stamens

stamens , ,45

45. 006-008

<Capsules>Colour

reddish46

46. 007-002

<Capsules>Endocarp

endocarp .47

47. 007-008

<Capsules>Hairs

glabrous48

48. 007-001

<Capsules>Length

3-5 mm long49 1.3-1.5 cm long50

49. 007-006, 50. 007-005

<Capsules>Locule number

bilocular (third locule rudimentary) or less often 3-locular51

51. 007-004

<Capsules>Pericarp

pericarp ;52

52. 007-007

<Capsules>Shape

with a cordiform outline, retuse at apex53

53. 007-003

<Flowers>Pedicels

pedicels .54

54. 005-001

<Habit>Height

3-12 m tall55

55. 001-001

<Leaves>Leaflets

leaflets , alternate, , , , ,56

56. 003-003

<Leaves>Petiole plus rachis

petiole plus rachis , slightly angled;57

57. 003-001

<Leaves>Petiolule

petiolules , , ;58

58. 003-002

<Leaves>Tertiary veins

tertiary veins reticulate.59

59. 003-005

<Leaves>Veins

venation prominent on abaxial surface, ;60

60. 003-004

<Seeds>Arillodes

nearly completely covered by a arillode61

61. 008-004

<Seeds>Colour

dark brown to black62

62. 008-001

<Seeds>Length

ca. 7 mm long63

63. 008-003

<Seeds>Shape

ellipsoid64

64. 008-002

<Stems>Hairs

puberulent or tomentulose, glabrescent with age65

65. 002-001

<Thyrses>Axes

axes , slightly angled.66

66. 004-004

<Thyrses>Length

10-25 cm long67

67. 004-002

<Thyrses>Position

axillary or distal68

68. 004-001

<Thyrses>Shape

paniculate69

69. 004-003

Calyx

Calyx , , , ,70

70. 006

Capsules

Capsule , , stipitate, , , (including a stipe);71

71. 007

Common Name

Creole (French Guiana): encens, kalima English (Suriname): basrakromantiehoedoe, gauetrie, kwassiewassitikie, se-gowtrie, warrimea, witte-gauetrie

Description

Tree 3-12 m tall. Stems puberulent or tomentulose, glabrescent with age. Leaves paripinnate; petiole plus rachis (5.5) 7-10 (20) cm long, slightly angled; petiolules 4-7 mm long, puberulent, enlarged at base; leaflets (4) 6 (8), alternate, chartaceous, glabrous, elliptic or oblong-elliptic, 8-17 (23) × 4-6.2 (7) cm, the base acute to obtuse, sometimes slightly asymmetrical, the apex obtuse or less often abruptly acuminate into an obtuse, retuse tip, the margins crenate; venation prominent on abaxial surface, usually drying darker than the blade; tertiary veins reticulate. Thyrses axillary or distal, 10-25 cm long, paniculate; axes tomentulose, slightly angled. Flowers in compound or simple dichasia; pedicels ca. 1 mm long. Calyx green, tomentulose, cup-shaped, 0.6-1 mm long, the lobes triangular, 0.4-0.5 mm long; petals white, rhombate, clawed, ca. 0.5 mm long, sparsely tomentose, with a single sericeous-tomentose appendage that is twice as long as the petal, and is fused to the petal’s base to form a pocket; disc puberulent; stamens 8, ca. 3 mm long, the filaments tomentose for most of their length; ovary appressed-pubescent. Capsule glabrous, reddish, stipitate, with a cordiform outline, retuse at apex, bilocular (third locule rudimentary) or less often 3-locular, 1.3-1.5 cm long (including a 3-5 mm long stipe); pericarp coriaceous; endocarp glabrous. Seed dark brown to black, ellipsoid, ca. 7 mm long, nearly completely covered by a white arillode.

Distribution

Guianas present, Southern America: Bolivia (Bolivia present); Colombia (Colombia present); Peru (Peru present); Venezuela (Venezuela present), Trinidad present

Widely distributed in Trinidad, Colombia, Venezuela, the Guianas, Brazil, Peru, and Bolivia (GU: 3; SU: 5; FG: 8).

Flowers

Flowers in compound or simple dichasia;72

72. 005

Habit

Tree .73

73. 001

Individuals Association

Guyana, Potaro-Siparuni, Pakaraima Mts., Ireng R., Henkel et al. 5469 Suriname, Para, Jodensavanne-Mapane Cr. area, Schulz 7276 Guyana, Upper Demerara-Berbice, Mabura Hill, Jansen-Jacobs et al. 1982 Bassin de l’Oyapock, Prévost & Grenand 2058 Bassin de l’Approuague, R. de l’Approuague, between Pierette and St. Grand Machieau, Oldeman 2519 Suriname, Brokopondo, van Blommesteijn lake, Tonka island, forest along beach, van Andel et al. 4667 Bassin de la Arataye, Sabatier 26

Leaves

Leaves paripinnate;74

74. 003

Notes

1 The label of the type collection at BM has been switched with that of Pseudima frutescens (Sapindus frutescens Aubl.).

Seeds

Seed , , , .75

75. 008

Stems

Stems .76

76. 002

Thyrses

Thyrses , , ;77

77. 004

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