Besleria penduliflora

<<<Calyx>Corolla>Base>Width

0.5 cm wide1

1. 005-006-004-001

<<<Calyx>Corolla>Limbs>Width

ca. 1.5 cm wide2

2. 005-006-006-001

<<<Calyx>Corolla>Lobes>Dimensions

0.4 x ca. 0.4 cm3

3. 005-006-008-002

<<<Calyx>Corolla>Lobes>Growth form

spreading4

4. 005-006-007-001

<<<Calyx>Corolla>Lobes>Shape

orbicular5

5. 005-006-008-001

<<<Calyx>Corolla>Margins>Margin type

entire6

6. 005-006-009-001

<<<Calyx>Corolla>Throat>Hairs

outside pilosulous, inside with a ring of glandular hairs in throat7

7. 005-006-005-002

<<<Calyx>Corolla>Throat>Width

1 cm wide8

8. 005-006-005-001

<<<Calyx>Corolla>Tube>Length

3-4 cm long9

9. 005-006-003-002

<<<Calyx>Corolla>Tube>Shape

cylindric10

10. 005-006-003-001

<<<Calyx>Ovary>Stigma>Lobe number

2-lobed11

11. 005-008-005-001

<<<Calyx>Ovary>Style>Hairs

densely strigillose12

12. 005-008-004-002

<<<Calyx>Ovary>Style>Length

3-3.5 cm long13

13. 005-008-004-001

<<<Leaves>Blade>Base>Hairs

above sparsely strigillose, below tomentose along veins14

14. 003-002-006-001

<<<Leaves>Blade>Larger blade>Dimensions

3.4-14.0 x 1.5-5.0 cm15

15. 003-002-003-001

<<<Leaves>Blade>Margins>Margin type

entire or remotely denticulate16

16. 003-002-004-001

<<Calyx>Apex>Growth form

recurved17

17. 005-005-001

<<Calyx>Apex>Hairs

outside strigillose, inside puberulous18

18. 005-005-002

<<Calyx>Corolla>Base

base with a spur, , middle not ventricose,19

19. 005-006-004

<<Calyx>Corolla>Colour

bright yellow20

20. 005-006-001

<<Calyx>Corolla>Length

3.5-4 cm long21

21. 005-006-002

<<Calyx>Corolla>Limbs

limb ,22

22. 005-006-006

<<Calyx>Corolla>Lobes

lobes reflexed, , ,23 lobes subequal, ,24

23. 005-006-008, 24. 005-006-007

<<Calyx>Corolla>Margins

margin ;25

25. 005-006-009

<<Calyx>Corolla>Throat

throat not constricted, , ,26

26. 005-006-005

<<Calyx>Corolla>Tube

tube , ,27

27. 005-006-003

<<Calyx>Margins>Margin type

entire28

28. 005-004-001

<<Calyx>Ovary>Dimensions

0.5 x 0.35 cm29

29. 005-008-002

<<Calyx>Ovary>Hairs

densely strigillose to pilosulous30

30. 005-008-003

<<Calyx>Ovary>Shape

ovoid31

31. 005-008-001

<<Calyx>Ovary>Stigma

stigma .32

32. 005-008-005

<<Calyx>Ovary>Style

style , ,33

33. 005-008-004

<<Calyx>Stamens>Relative distance

inserted ca. 0.8 cm above corolla base34

34. 005-007-001

<<Calyx>Tube>Dimensions

0.4-0.7 x 0.2-0.4 cm35

35. 005-003-004

<<Calyx>Tube>Length

ca. 0.1 cm long36

36. 005-003-001

<<Calyx>Tube>Lobe number

437

37. 005-003-002

<<Calyx>Tube>Shape

all lanceolate to oblong38

38. 005-003-003

<<Flowers>Pedicels>Hairs

subtomentose39

39. 004-003-002

<<Flowers>Pedicels>Length

1.5-2 cm long40

40. 004-003-001

<<Flowers>Peduncle>Hairs

tomentose41

41. 004-002-002

<<Flowers>Peduncle>Length

(2.5-) 5-7 cm long42

42. 004-002-001

<<Leaves>Blade>Apex

apex acuminate,43

43. 003-002-005

<<Leaves>Blade>Base

base cuneate, .44

44. 003-002-006

<<Leaves>Blade>Larger blade

larger blade45

45. 003-002-003

<<Leaves>Blade>Margins

margin ,46

46. 003-002-004

<<Leaves>Blade>Shape

lanceolate to elliptic or oblanceolate47

47. 003-002-002

<<Leaves>Blade>Texture

papyraceous when dry48

48. 003-002-001

<<Leaves>Petiole>Hairs

tomentose49

49. 003-001-002

<<Leaves>Petiole>Length

0.5-3.2 cm long50

50. 003-001-001

<<Stems>Internodes>Hairs

tomentose to glabrescent51

51. 002-002-001

<Calyx>Apex

apex acute and , ;52

52. 005-005

<Calyx>Colour

greenish-yellow53

53. 005-001

<Calyx>Corolla

corolla strongly oblique in calyx, , ,54

54. 005-006

<Calyx>Lobes

lobes nearly free,55

55. 005-002

<Calyx>Margins

margin ,56

56. 005-004

<Calyx>Ovary

ovary , , ,57

57. 005-008

<Calyx>Stamens

stamens included or slightly exserted, ;58

58. 005-007

<Calyx>Tube

tube , lobes subequal, with dorsal lobe curved around corolla spur and narrower, , ,59

59. 005-003

<Flowers>Flower number per inflorescence

1-3-flowered60

60. 004-001

<Flowers>Pedicels

pedicel ,61

61. 004-003

<Flowers>Peduncle

peduncle , ;62

62. 004-002

<Habit>Height

1.5-6 m tall63

63. 001-001

<Leaves>Blade

blade , , ,64

64. 003-002

<Leaves>Petiole

petiole , ;65

65. 003-001

<Mature berries>Colour

greenish with red pericarp66

66. 006-001

<Mature berries>Dimensions

1-1.1 x 1-1.1 cm67

67. 006-003

<Mature berries>Shape

globose68

68. 006-002

<Stems>Growth form

erect69

69. 002-001

<Stems>Internodes

internodes .70

70. 002-002

Calyx

Calyx subcampanulate, ,71

71. 005

Description

Terrestrial shrub, 1.5-6 m tall. Stem woody, erect, internodes tomentose to glabrescent. Leaves unequal in a pair; petiole 0.5-3.2 cm long, tomentose; blade papyraceous when dry, lanceolate to elliptic or oblanceolate, larger blade 3.4-14.0 x 1.5-5.0 cm, margin entire or remotely denticulate, apex acuminate, base cuneate, above sparsely strigillose, below tomentose along veins. Flowers in umbellate or subcymose, 1-3-flowered inflorescences; peduncle (2.5-) 5-7 cm long, tomentose; pedicel 1.5-2 cm long, subtomentoseCalyx subcampanulate, greenish-yellow, lobes nearly free, tube ca. 0.1 cm long, 4 lobes subequal, with dorsal lobe curved around corolla spur and narrower, all lanceolate to oblong, 0.4-0.7 x 0.2-0.4 cm, margin entire, apex acute and recurved, outside strigillose, inside puberulous; corolla strongly oblique in calyx, bright yellow, 3.5-4 cm long, tube cylindric, 3-4 cm long, base with a spur, 0.5 cm wide, middle not ventricose, throat not constricted, 1 cm wide, outside pilosulous, inside with a ring of glandular hairs in throat, limb ca. 1.5 cm wide, lobes subequal, spreading, lobes reflexed, orbicular, 0.4 x ca. 0.4 cm, margin entire; stamens included or slightly exserted, inserted ca. 0.8 cm above corolla base; ovary ovoid, 0.5 x 0.35 cm, densely strigillose to pilosulous, style 3-3.5 cm long, densely strigillose, stigma 2-lobed. Mature berry greenish with red pericarp, globose, 1-1.1 x 1-1.1 cm.

Distribution

Guayana Highlands present, Guyana present, Southern America: Venezuela (Venezuela present)

Guayana highlands of Venezuela and Guyana; 16 specimens studied (GU: 7).

Flowers

Flowers in umbellate or subcymose, inflorescences;72

72. 004

Habit

Terrestrial shrub, .73

73. 001

Individuals Association

Guyana, Cuyuni-Mazaruni Region, NW of northern prow of Mt. Roraima, Hahn et al. 5468 Guyana, Potaro-Siparuni Region, upper Potaro River Region, upper slopes of Mt. Wokomung, Boom & Samuels 9150 Guyana, upper Mazaruni R. basin, Mt. Ayanganna, Tillett & Tillett 45190 Guyana, Mt. Kopinang, 1350 m, Hahn et al. 4299

Leaves

Leaves unequal in a pair;74

74. 003

Mature berries

Mature berry , , .75

75. 006

Notes

Besleria penduliflora resembles and is probably closely related to B. pendula Hanst. from Venezuela but the latter species differs in having corollas only 1.7-2.4 cm long, calyx lobes pilosulous only at base, and pubescent leaves.

Phenology

Collected in flower in ; fruiting .

Stems

Stem woody, ,76

76. 002

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