Besleria saxicola

<<<Calyx>Corolla>Base>Width

0.3-0.5 cm wide1

1. 005-005-004-001

<<<Calyx>Corolla>Limbs>Width

0.7-1 cm wide2

2. 005-005-006-001

<<<Calyx>Corolla>Lobes>Dimensions

0.15-0.2 x 0.15-0.2 cm3

3. 005-005-007-003

<<<Calyx>Corolla>Lobes>Growth form

slightly spreading4

4. 005-005-007-001

<<<Calyx>Corolla>Lobes>Shape

orbicular5

5. 005-005-007-002

<<<Calyx>Corolla>Margins>Margin type

entire6

6. 005-005-008-001

<<<Calyx>Corolla>Throat>Hairs

outside glabrous, inside with a ring of glandular hairs in throat7

7. 005-005-005-002

<<<Calyx>Corolla>Throat>Width

0.5-0.6 cm wide8

8. 005-005-005-001

<<<Calyx>Corolla>Tube>Length

1.7-1.8 cm long9

9. 005-005-003-002

<<<Calyx>Corolla>Tube>Shape

cylindric10

10. 005-005-003-001

<<<Calyx>Ovary>Stigma>Lobe number

2-lobed11

11. 005-007-005-001

<<<Calyx>Ovary>Style>Hairs

puberulous12

12. 005-007-004-002

<<<Calyx>Ovary>Style>Length

ca. 1-1.5 cm long13

13. 005-007-004-001

<<<Leaves>Blade>Base>Hairs

above strigillose, below hirsute especially along midrib and veins14

14. 003-002-006-001

<<<Leaves>Blade>Margins>Margin type

remotely sharp-serrulate15

15. 003-002-004-001

<<Calyx>Apex>Hairs

outside hirsute, inside glabrous16

16. 005-004-001

<<Calyx>Corolla>Base

base gibbous, , middle somewhat ventricose,17

17. 005-005-004

<<Calyx>Corolla>Colour

yellow18

18. 005-005-001

<<Calyx>Corolla>Length

1.8-2 cm long19

19. 005-005-002

<<Calyx>Corolla>Limbs

limb ,20

20. 005-005-006

<<Calyx>Corolla>Lobes

lobes subequal, , , ,21

21. 005-005-007

<<Calyx>Corolla>Margins

margin ;22

22. 005-005-008

<<Calyx>Corolla>Throat

throat constricted, , ,23

23. 005-005-005

<<Calyx>Corolla>Tube

tube , ,24

24. 005-005-003

<<Calyx>Lobes>Dimensions

0.6-0.8 x 0.2-0.3 cm25

25. 005-002-003

<<Calyx>Lobes>Growth form

erect26

26. 005-002-001

<<Calyx>Lobes>Shape

ovate to ovate-lanceolate27

27. 005-002-002

<<Calyx>Margins>Margin type

entire and long-ciliate28

28. 005-003-001

<<Calyx>Ovary>Dimensions

0.2-0.3 x 0.2-0.3 cm29

29. 005-007-002

<<Calyx>Ovary>Hairs

glabrous30

30. 005-007-003

<<Calyx>Ovary>Shape

conic31

31. 005-007-001

<<Calyx>Ovary>Stigma

stigma .32

32. 005-007-005

<<Calyx>Ovary>Style

style , ,33

33. 005-007-004

<<Flowers>Pedicels>Hairs

hirsute34

34. 004-002-002

<<Flowers>Pedicels>Length

(0.5-)1-1.5 cm long35

35. 004-002-001

<<Leaves>Blade>Apex

apex short-acuminate,36

36. 003-002-005

<<Leaves>Blade>Base

base cuneate, .37

37. 003-002-006

<<Leaves>Blade>Dimensions

20-30 x 7-13 cm38

38. 003-002-003

<<Leaves>Blade>Margins

margin ,39

39. 003-002-004

<<Leaves>Blade>Shape

obliquely elliptic40

40. 003-002-002

<<Leaves>Blade>Texture

membranous when dry41

41. 003-002-001

<<Leaves>Petiole>Hairs

hirsute42

42. 003-001-002

<<Leaves>Petiole>Length

3-12 cm long43

43. 003-001-001

<<Stems>Apex>Hairs

hirsute, glabrescent below44

44. 002-002-001

<Calyx>Apex

apex subulate-acuminate, ;45

45. 005-004

<Calyx>Colour

yellowish46

46. 005-001

<Calyx>Corolla

corolla oblique in calyx, , ,47

47. 005-005

<Calyx>Lobes

lobes free, , subequal, , ,48

48. 005-002

<Calyx>Margins

margin ,49

49. 005-003

<Calyx>Ovary

ovary , , ,50

50. 005-007

<Calyx>Stamens

stamens included, inserted below middle of corolla tube;51

51. 005-006

<Flowers>Flower number per inflorescence

many-flowered52

52. 004-001

<Flowers>Pedicels

pedicel , .53

53. 004-002

<Habit>Height

0.3-1.8 m tall54

54. 001-001

<Leaves>Blade

blade becoming , , ,55

55. 003-002

<Leaves>Petiole

petiole , ;56

56. 003-001

<Mature berries>Colour

yellow57

57. 006-001

<Mature berries>Dimensions

0.8-1 x 0.8-1 cm58

58. 006-003

<Mature berries>Shape

globose59

59. 006-002

<Stems>Apex

apex .60

60. 002-002

<Stems>Growth form

erect61

61. 002-001

Calyx

Calyx subcampanulate, ,62

62. 005

Description

Terrestrial herb, subshrub or shrub, 0.3-1.8 m tall. Stem woody at base, erect, apex hirsute, glabrescent below. Leaves equal to subequal in a pair; petiole 3-12 cm long, hirsute; blade becoming membranous when dry, obliquely elliptic, 20-30 x 7-13 cm, margin remotely sharp-serrulate, apex short-acuminate, base cuneate, above strigillose, below hirsute especially along midrib and veins. Flowers in fasciculate many-flowered, epedunculate inflorescences; pedicel (0.5-)1-1.5 cm long, hirsute. Calyx subcampanulate, yellowish, lobes free, erect, subequal, ovate to ovate-lanceolate, 0.6-0.8 x 0.2-0.3 cm, margin entire and long-ciliate, apex subulate-acuminate, outside hirsute, inside glabrous; corolla oblique in calyx, yellow, 1.8-2 cm long, tube cylindric, 1.7-1.8 cm long, base gibbous, 0.3-0.5 cm wide, middle somewhat ventricose, throat constricted, 0.5-0.6 cm wide, outside glabrous, inside with a ring of glandular hairs in throat, limb 0.7-1 cm wide, lobes subequal, slightly spreading, orbicular, 0.15-0.2 x 0.15-0.2 cm, margin entire; stamens included, inserted below middle of corolla tube; ovary conic, 0.2-0.3 x 0.2-0.3 cm, glabrous, style ca. 1-1.5 cm long, puberulous, stigma 2-lobed. Mature berry yellow, globose, 0.8-1 x 0.8-1 cm.

Distribution

Guyana endemic

Endemic to Guyana; 11 specimens studied (GU: 11).

Flowers

Flowers in fasciculate , epedunculate inflorescences;63

63. 004

Habit

Terrestrial herb, subshrub or shrub, .64

64. 001

Individuals Association

Guyana, Cuyuni-Mazaruni Region, Eping R., McDowell et al. 3851 Guyana, Essequibo R., Moraballi Cr., near Bartica, Sandwith 58 Guyana, Potaro-Siparuni Region, Tumatumari, Gleason 420 Guyana, Barima-Waini Region, between Aranka Head and Barima Head, NW of Kariako R., McDowell et al. 4379 Guyana, Cuyuni-Mazaruni Region, Kuwara, Kurupung R., Altson 311

Leaves

Leaves equal to subequal in a pair;65

65. 003

Mature berries

Mature berry , , .66

66. 006

Phenology

Flowering in , and fruiting in .

Stems

Stem woody at base, ,67

67. 002

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