Talisia squarrosa

<<<Calyx>Ovary>Stigma>Shape

elongated, trigonous1

1. 005-008-003-001

<<<Calyx>Petals>Apex>Shape

obtuse2

2. 005-004-006-001

<<<Calyx>Petals>Base>Shape

cuneate to nearly clawed3

3. 005-004-005-001

<<<Calyx>Stamens>Anthers>Length

0.7-0.8 mm long4

4. 005-007-003-002

<<<Calyx>Stamens>Anthers>Shape

ovate, obtuse at apex5

5. 005-007-003-001

<<<Calyx>Stamens>Filaments>Hairs

glabrous6

6. 005-007-002-001

<<<Calyx>Stamens>Filaments>Length

1-2 mm long7

7. 005-007-002-002

<<<Habit>Bark>Inner bark>Colour

reddish brown8

8. 001-003-002-001

<<<Leaves>Leaflets>Apex>Shape

obtuse or acute9

9. 003-003-009-001

<<<Leaves>Leaflets>Base>Shape

acute or obtuse, asymmetrical10

10. 003-003-010-001

<<Calyx>Appendages>Hairs

adaxially tomentose11

11. 005-005-002

<<Calyx>Appendages>Shape

deltate12

12. 005-005-001

<<Calyx>Disk>Hairs

minutely tomentose13

13. 005-006-003

<<Calyx>Disk>Height

0.6-0.7 mm tall14

14. 005-006-004

<<Calyx>Disk>Lobe number

5-lobed15

15. 005-006-002

<<Calyx>Disk>Shape

cup-shaped16

16. 005-006-001

<<Calyx>Ovary>Hairs

sericeous-tomentose17

17. 005-008-002

<<Calyx>Ovary>Shape

conical18

18. 005-008-001

<<Calyx>Ovary>Stigma

the stigma .19

19. 005-008-003

<<Calyx>Petals>Apex

the apex ;20

20. 005-004-006

<<Calyx>Petals>Base

the base ,21

21. 005-004-005

<<Calyx>Petals>Colour

white22

22. 005-004-001

<<Calyx>Petals>Hairs

abaxially appressed-pubescent on lower half23

23. 005-004-004

<<Calyx>Petals>Length

2.8-3 mm long24

24. 005-004-003

<<Calyx>Petals>Shape

rhombo-elliptic or wide elliptic25

25. 005-004-002

<<Calyx>Sepals>Shape

elliptic to ovate, concave26

26. 005-003-001

<<Calyx>Stamens>Anthers

the anthers , ;27

27. 005-007-003

<<Calyx>Stamens>Filaments

the filaments , of unequal length, ,28

28. 005-007-002

<<Calyx>Stamens>Stamen number

829

29. 005-007-001

<<Habit>Bark>Colour

silvery gray30

30. 001-003-001

<<Habit>Bark>Inner bark

inner bark31

31. 001-003-002

<<Habit>Trunk>Buttresses

buttressed at base32

32. 001-002-002

<<Habit>Trunk>Diameter

reaching 30 cm in diam.33

33. 001-002-001

<<Immature fruits>Pericarp>Texture

woody, smooth34

34. 006-003-001

<<Leaves>Leaflets>Abaxial surfaces

the abaxial surface with prominent primary and secondary veins,35

35. 003-003-008

<<Leaves>Leaflets>Adaxial surfaces

the adaxial surface with prominulous venation,36

36. 003-003-007

<<Leaves>Leaflets>Apex

the apex ,37

37. 003-003-009

<<Leaves>Leaflets>Base

the base ;38

38. 003-003-010

<<Leaves>Leaflets>Colour

dull39

39. 003-003-004

<<Leaves>Leaflets>Dimensions

5.3-9(19) × 2.4-4.5(7.5) cm40

40. 003-003-006

<<Leaves>Leaflets>Hairs

glabrous41

41. 003-003-003

<<Leaves>Leaflets>Leaflet number

242

42. 003-003-001

<<Leaves>Leaflets>Shape

elliptic43

43. 003-003-005

<<Leaves>Leaflets>Texture

coriaceous44

44. 003-003-002

<<Leaves>Petiole>Hairs

glabrous45

45. 003-001-002

<<Leaves>Petiole>Length

2-5.5 cm long46

46. 003-001-001

<<Leaves>Petiole>Lenticels

lenticellate47

47. 003-001-004

<<Leaves>Petiole>Shape

flattened along adaxial surface48

48. 003-001-003

<<Leaves>Petiolule>Colour

drying dark brown49

49. 003-002-003

<<Leaves>Petiolule>Hairs

glabrous50

50. 003-002-001

<<Leaves>Petiolule>Length

3-7 mm long51

51. 003-002-002

<<Seeds>Sarcotesta>Colour

white52

52. 007-002-001

<<Thyrses>Axes>Hairs

puberulent53

53. 004-003-001

<<Thyrses>Bracts and bracteoles>Length

< 1 mm long54

54. 004-004-002

<<Thyrses>Bracts and bracteoles>Shape

subulate55

55. 004-004-001

<<Thyrses>Pedicels>Hairs

tomentulose56

56. 004-007-002

<<Thyrses>Pedicels>Length

2-3 mm long57

57. 004-007-001

<<Thyrses>Peduncle>Hairs

puberulent or minutely tomentose58

58. 004-006-002

<<Thyrses>Peduncle>Length

2-4 mm long59

59. 004-006-001

<Branches>Hairs

glabrous60

60. 002-003

<Branches>Lenticels

lenticellate61

61. 002-002

<Branches>Shape

terete62

62. 002-001

<Calyx>Appendages

appendages as long as the petal, , as wide as the petal or narrower, abaxially papillate, ;63

63. 005-005

<Calyx>Disk

disc , , , ;64

64. 005-006

<Calyx>Hairs

tomentose65

65. 005-002

<Calyx>Length

3-4 mm long66

66. 005-001

<Calyx>Ovary

ovary , ,67

67. 005-008

<Calyx>Petals

petals , , , adaxially minutely papillate, ,68

68. 005-004

<Calyx>Sepals

the sepals free to base, ;69

69. 005-003

<Calyx>Stamens

stamens ,70

70. 005-007

<Habit>Bark

bark , .71

71. 001-003

<Habit>Height

to 20-30 m tall72

72. 001-001

<Habit>Trunk

trunk , ;73

73. 001-002

<Immature fruits>Hairs

sericeous-tomentose74

74. 006-002

<Immature fruits>Pericarp

the pericarp .75

75. 006-003

<Immature fruits>Shape

ellipsoid or ovoid76

76. 006-001

<Leaves>Leaflets

leaflets , opposite, , , , , ,77

77. 003-003

<Leaves>Petiole

petiole , , , ;78

78. 003-001

<Leaves>Petiolule

petiolules pulvinate, , , ;79

79. 003-002

<Leaves>Tertiary veins

tertiary veins reticulate, prominulous.80

80. 003-004

<Seeds>Sarcotesta

with juicy sarcotesta (fide Polak 1992)81

81. 007-002

<Seeds>Shape

ellipsoid82

82. 007-001

<Thyrses>Axes

axes obtusely angled, sulcate, ;83

83. 004-003

<Thyrses>Bracts and bracteoles

bracts and bracteoles , , early deciduous;84

84. 004-004

<Thyrses>Dichasia

dichasia compound;85

85. 004-005

<Thyrses>Length

to 27 cm long86

86. 004-001

<Thyrses>Pedicels

pedicels , , articulate at the middle.87

87. 004-007

<Thyrses>Peduncle

peduncles , ;88

88. 004-006

<Thyrses>Position

distal or axillary on distal portion of branches89

89. 004-002

Branches

Branches , rough, , .90

90. 002

Calyx

Calyx , ,91

91. 005

Common Name

English (French Guiana): grigri, singabassou English (Guyana): candlewood, karimora,, morabali, muro-balli, sand mora

Description

Tree to 20-30 m tall; trunk reaching 30 cm in diam., buttressed at base; bark silvery gray, inner bark reddish brown. Branches terete, rough, lenticellate, glabrous. Leaves paripinnate; petiole 2-5.5 cm long, glabrous, flattened along adaxial surface, lenticellate; petiolules pulvinate, glabrous, 3-7 mm long, drying dark brown; leaflets 2, opposite, coriaceous, glabrous, dull, elliptic, 5.3-9(19) × 2.4-4.5(7.5) cm, the adaxial surface with prominulous venation, the abaxial surface with prominent primary and secondary veins, the apex obtuse or acute, the base acute or obtuse, asymmetrical; tertiary veins reticulate, prominulous. Thyrses panicle-shaped, to 27 cm long, distal or axillary on distal portion of branches; axes obtusely angled, sulcate, puberulent; bracts and bracteoles subulate, < 1 mm long, early deciduous; dichasia compound; peduncles 2-4 mm long, puberulent or minutely tomentose; pedicels 2-3 mm long, tomentulose, articulate at the middle. Calyx 3-4 mm long, tomentose, the sepals free to base, elliptic to ovate, concave; petals white, rhombo-elliptic or wide elliptic, 2.8-3 mm long, adaxially minutely papillate, abaxially appressed-pubescent on lower half, the base cuneate to nearly clawed, the apex obtuse; appendages as long as the petal, deltate, as wide as the petal or narrower, abaxially papillate, adaxially tomentose; disc cup-shaped, 5-lobed, minutely tomentose, 0.6-0.7 mm tall; stamens 8, the filaments glabrous, of unequal length, 1-2 mm long, the anthers ovate, obtuse at apex, 0.7-0.8 mm long; ovary conical, sericeous-tomentose, the stigma elongated, trigonous. Fruits (immature) ellipsoid or ovoid, sericeous-tomentose, the pericarp woody, smooth. Seed ellipsoid, with white juicy sarcotesta (fide Polak 1992).

Distribution

French Guiana present, Guyana present, Suriname present

Known from Guyana, Suriname, and French Guiana (GU: 10; SU: 2; FG: 4).

Habit

Tree ;92

92. 001

Immature fruits

Fruits (immature) , ,93

93. 006

Individuals Association

Organabo Station, forest on white sands, Loubry 2382 Guyana, Berbice, Mora-mora-bisi Cr., Courantyne R., white sand, Hohenkerk 715 Region Littorale, Cayenne-St. Laurent road, km 171.5, Sastre et al. 8222 Organabo, forest on white sands ca. 2 km W of Organabo R., north of road, Acevedo-Rdgz. & Elfort 11117 Suriname, Sipalawini, Summit of Tafelberg, Evans 3202 Guyana, Demerara R., sandhills, Wallaba forest, Fanshawe 1215 Guyana, Barima, Amakura R., de la Cruz 3475

Leaves

Leaves paripinnate;94

94. 003

Seeds

Seed , .95

95. 007

Thyrses

Thyrses panicle-shaped, , ;96

96. 004

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